H. sapho flies in the primary forest along with its comimic H. cydno. H. sapho is rare at La Selva, like its obligate host P. pittieri. It is a specialized group feeder, like H. doris or H. sara. Larvae look very much like those of H. sara, except that 5th instar larvae are yellow dorsally instead of black. Like H. cydno, H. sapho has a completely different color form on the southwest side of Costa Rica, called H. hewitsoni. Range: Mexico to Colombia and Ecuador.

H. hewitsoni has been studied extensively in Corcovado National Park. One interesting finding, according to Erika Deinert, is that H. hewitsoni larvae can survive, if poorly, on P. arbelaezii from La Selva. Feeding trials with Red Pedilia, the P. pittieri specialist flea beetle, show the same pattern: limited feeding and growth on P. arbelaezii and P. biflora.